UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F COVER PAGE
OMB APPROVAL
OMB Number:3235-0006
Expires:July 31, 2015
Estimated average burden
hours per response:23.8
Report for the Calendar Year or Quarter Ended:
09-30-2022
Check here if Amendment: ☐ Amendment Number:
This Amendment (Check only one.):☐ is a restatement.
☐ adds new holdings entries.

Institutional Investment Manager Filing this Report:

Name:
Granahan Investment Management, LLC
Address:
404 WYMAN STREET
SUITE 460
WALTHAM, MA 02451
Form 13F File Number:
028-05876

The institutional investment manager filing this report and the person by whom it is signed hereby represent that the person signing the report is authorized to submit it, that all information contained herein is true, correct and complete, and that it is understood that all required items, statements, schedules, lists, and tables, are considered integral parts of this form.


Person Signing this Report on Behalf of Reporting Manager:
Name:
Susan Rawley
Title:
VP/Operations and Account Administration
Phone:
781-902-1413


Signature, Place, and Date of Signing:

Susan Rawley
                         [Signature]
Waltham, MASSACHUSETTS
                 [City, State]
11-14-2022
                           [Date]
                          

Report Type (Check only one.):

☒ 13F HOLDINGS REPORT. (Check here if all holdings of this reporting manager are reported in this report.) 

☐ 13F NOTICE. (Check here if no holdings reported are in this report, and all holdings are reported by other reporting manager(s).) 

☐ 13F COMBINATION REPORT. (Check here if a portion of the holdings for this reporting manager are reported in this report and a portion are reported by other reporting manager(s).)


Form 13F Summary Page

Report Summary:

Number of Other Included Managers:
1
Form 13F Information table Entry Total:
176
Form 13F Information table Value Total:
3062803
          (thousands)

List of Other Included Managers:

Provide a numbered list of the name(s) and Form 13F file number(s) of all institutional investment managers with respect to which this report is filed, other than the manager filing this report.
[If there are no entries in this list, state "NONE" and omit the column headings and list entries.]

No.Form 13F File NumberNameCIK
1
028-15699
Destra Capital Advisors LLC
 


UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE
OMB APPROVAL
OMB Number:3235-0006
Expires:July 31, 2015
Estimated average burden
hours per response:23.8
COLUMN 1COLUMN 2COLUMN 3COLUMN 4COLUMN 5COLUMN 6COLUMN 7COLUMN 8
VALUESHRS ORSH/PUT/INVESTMENTOTHERVOTING AUTHORITY
NAME OF ISSUERTITLE OF CLASSCUSIP(x$1000)PRN AMTPRNCALLDISCRETIONMANAGERSOLESHAREDNONE
908 DEVICES INCCOM65443P10215,913967,364SHSOLE819,440 0 147,924
AFFIMED N VCOMN010451087,9123,840,990SHSOLE3,646,810 0 194,180
AIRGAIN INCCOM00938A1044,006555,677SHSOLE446,114 0 109,563
AKOUSTIS TECHNOLOGIES INCCOM00973N1022,283768,810SHSOLE605,900 0 162,910
AKOYA BIOSCIENCES INCCOM00974H1044,686398,824SHSOLE348,503 0 50,321
ALTA EQUIPMENT GROUP INCCOMMON STOCK02128L10610,965995,880SHSOLE874,147 0 121,733
AMBARELLA INCSHSG037AX10110,848193,101SHSOLE180,782 0 12,319
AMERESCO INCCL A02361E10818,912284,472SHSOLE273,180 0 11,292
AMERICAN WELL CORPCL A03044L1055,9941,669,539SHSOLE1,440,638 0 228,901
APELLIS PHARMACEUTICALS INCCOM03753U1069,339136,735SHSOLE110,459 0 26,276
APPIAN CORPCL A03782L1013,14677,054SHSOLE63,855 0 13,199
ARCUS BIOSCIENCES INCCOM03969F1098,982343,335SHSOLE285,362 0 57,973
ARRAY TECHNOLOGIES INCCOM SHS04271T10065,3143,939,313SHSOLE3,743,463 0 195,850
ARTERIS INCCOM04302A1042,166325,166SHSOLE255,657 0 69,509
ATRICURE INCCOM04963C2097,101181,602SHSOLE146,090 0 35,512
AXON ENTERPRISE INCCOM05464C101104,849905,824SHSOLE740,328 0 165,496
AXT INCCOM00246W1036,8071,015,961SHSOLE836,211 0 179,750
AZENTA INCCOM11434010248,3001,126,917SHSOLE919,988 0 206,929
BEAM THERAPEUTICS INCCOM07373V1055,261110,435SHSOLE92,456 0 17,979
BIOLIFE SOLUTIONS INCCOM NEW09062W20416,832739,873SHSOLE634,973 0 104,900
BIOMERICA INCCOM NEW09061H3074,4001,119,687SHSOLE1,026,950 0 92,737
BLUEBIRD BIO INCCOM09609G1003,729589,083SHSOLE514,306 0 74,777
BLUEPRINT MEDICINES CORPCOM09627Y1099,500144,177SHSOLE116,467 0 27,710
BUMBLE INCCOM CL A12047B10518,569864,077SHSOLE709,111 0 154,966
CARIBOU BIOSCIENCES INCCOM1420381082,513238,204SHSOLE215,904 0 22,300
CASELLA WASTE SYS INCCL A14744810422,517294,765SHSOLE239,675 0 55,090
CASTLE BIOSCIENCES INCCOM14843C10519,879762,217SHSOLE663,024 0 99,193
CEVA INCCOM15721010510,839413,232SHSOLE352,790 0 60,442
CHART INDS INCCOM16115Q30845,522246,934SHSOLE226,247 0 20,687
CHEGG INCCOM16309210920,427969,497SHSOLE794,052 0 175,445
CHICKEN SOUP FOR THE SOUL ENCL A16842Q1008,8611,287,965SHSOLE1,229,416 0 58,549
CIENA CORPCOM NEW1717793096,754167,051SHSOLE135,254 0 31,797
CLEAN ENERGY FUELS CORPCOM1844991017,8161,463,728SHSOLE1,376,945 0 86,783
COLUMBUS MCKINNON CORP N YCOM19933310513,547517,867SHSOLE447,624 0 70,243
COMPUTER TASK GROUP INCCOM2054771021,214181,163SHSOLE181,163 0 0
CONFLUENT INCCLASS A COM20717M10361625,903SHSOLE20,338 0 5,565
CORVEL CORPCOM2210061095,25537,959SHSOLE30,777 0 7,182
COSTAR GROUP INCCOM22160N109105,3271,512,227SHSOLE1,249,613 0 262,614
COUPA SOFTWARE INCCOM22266L1068,960152,380SHSOLE124,990 0 27,390
CRISPR THERAPEUTICS AGNAMEN AKTH171821083,39952,006SHSOLE42,514 0 9,492
CRYOPORT INCCOM PAR $0.00122905030711,691479,908SHSOLE438,946 0 40,962
CUSTOMERS BANCORP INCCOM23204G1002,92899,311SHSOLE91,481 0 7,830
CYBERARK SOFTWARE LTDSHSM2682V1083,92826,196SHSOLE20,630 0 5,566
CYTOSORBENTS CORPCOM NEW23283X2062,0551,511,029SHSOLE1,351,219 0 159,810
DARLING INGREDIENTS INCCOM2372661016,09292,094SHSOLE92,094 0 0
DEFINITIVE HEALTHCARE CORPCLASS A COM24477E10312,789823,003SHSOLE705,574 0 117,429
DERMTECH INCCOM24984K105836211,185SHSOLE211,185 0 0
DIGITAL TURBINE INCCOM NEW25400W10246,0753,197,441SHSOLE3,021,916 0 175,525
DISTRIBUTION SOLUTIONS GRP ICOM52077610550617,951SHSOLE17,951 0 0
DOMO INCCOM CL B25755410513,060725,937SHSOLE600,119 0 125,818
DOUGLAS DYNAMICS INCCOM25960R10573426,205SHSOLE26,205 0 0
EDGIO INCCOM53261M10425,5379,185,946SHSOLE8,169,496 0 1,016,450
ENOVIX CORPORATIONCOM29359410731,7071,729,297SHSOLE1,429,422 0 299,875
ENPHASE ENERGY INCCOM29355A107114,779413,662SHSOLE352,946 0 60,716
EPAM SYS INCCOM29414B1046,70318,506SHSOLE18,506 0 0
ETSY INCCOM29786A10677,080769,799SHSOLE639,621 0 130,178
EURONET WORLDWIDE INCCOM29873610937,399493,649SHSOLE459,567 0 34,082
EVERBRIDGE INCCOM29978A1042,51681,470SHSOLE75,211 0 6,259
EVERQUOTE INCCOM CL A30041R10811,1721,638,103SHSOLE1,359,149 0 278,954
EVOLENT HEALTH INCCL A30050B10188,8962,474,135SHSOLE2,179,513 0 294,622
EXPEDIA GROUP INCCOM NEW30212P3031,87620,021SHSOLE20,021 0 0
FATE THERAPEUTICS INCCOM31189P1027,315326,419SHSOLE264,643 0 61,776
FIGS INCCL A30260D10315,0151,819,973SHSOLE1,734,405 0 85,568
FIRST SOLAR INCCOM33643310721,090159,446SHSOLE148,775 0 10,671
FIRST WATCH RESTAURANT GROUPCOM33748L10113,327920,356SHSOLE750,833 0 169,523
GENIUS SPORTS LIMITEDSHARES CL AG3934V10933,9719,256,521SHSOLE7,683,520 0 1,573,001
GENTHERM INCCOM37253A10312,312247,575SHSOLE236,866 0 10,709
GLOBANT S ACOML4438510929,294156,588SHSOLE128,637 0 27,951
GREEN BRICK PARTNERS INCCOM3927091014,785223,828SHSOLE198,857 0 24,971
GRID DYNAMICS HLDGS INCCL A39813G10915,933850,684SHSOLE813,318 0 37,366
GROCERY OUTLET HLDG CORPCOM39874R1014,090122,858SHSOLE100,447 0 22,411
GUIDEWIRE SOFTWARE INCCOM40171V1008,972145,698SHSOLE116,261 0 29,437
HAGERTY INCCL A COM40516610912,4671,386,718SHSOLE1,325,183 0 61,535
HARMONIC INCCOM41316010214,0751,076,911SHSOLE1,013,068 0 63,843
HARVARD BIOSCIENCE INCCOM4169061053,8231,493,269SHSOLE1,415,987 0 77,282
HEALTH CATALYST INCCOM42225T1076,289648,299SHSOLE550,571 0 97,728
HEALTHEQUITY INCCOM42226A10746,493692,166SHSOLE565,435 0 126,731
HESKA CORPCOM RESTRC NEW42805E3064,20457,654SHSOLE46,775 0 10,879
HUBSPOT INCCOM44357310041,618154,072SHSOLE126,347 0 27,725
HUDSON TECHNOLOGIES INCCOM4441441097,229983,527SHSOLE907,354 0 76,173
ICAD INCCOM NEW44934S2065,3082,527,839SHSOLE2,397,257 0 130,582
IMMUNOGEN INCCOM45253H1018,9721,876,960SHSOLE1,569,843 0 307,117
INDIE SEMICONDUCTOR INCCLASS A COM45569U10133,0714,517,864SHSOLE4,324,114 0 193,750
INMODE LTDSHSM5425M1034,587157,571SHSOLE151,599 0 5,972
INNOVATIVE INDL PPTYS INCCOM45781V1013,84443,430SHSOLE36,278 0 7,152
INTELLIA THERAPEUTICS INCCOM45826J1058,441150,835SHSOLE124,751 0 26,084
ISHARES TRRUS 2000 GRW ETF4642876481,1965,788SHSOLE5,788 0 0
JAMES RIV GROUP LTDCOMG5005R10722,189972,755SHSOLE810,683 0 162,072
KEZAR LIFE SCIENCES INCCOM49372L1003,852447,418SHSOLE388,111 0 59,307
KINIKSA PHARMACEUTICALS LTDCOM CL AG5269C1017,092552,350SHSOLE466,732 0 85,618
KINSALE CAP GROUP INCCOM49714P1086,73426,363SHSOLE21,334 0 5,029
KORNIT DIGITAL LTDSHSM6372Q11387,7553,297,827SHSOLE2,933,785 0 364,042
KULICKE & SOFFA INDS INCCOM50124210120,525532,711SHSOLE510,046 0 22,665
LENDINGTREE INC NEWCOM52603B10713,222554,145SHSOLE513,086 0 41,059
LIFE TIME GROUP HOLDINGS INCCOMMON STOCK53190C10216,9301,736,395SHSOLE1,422,934 0 313,461
LIFEMD INCCOM53216B10419096,495SHSOLE96,495 0 0
LIQUIDITY SVCS INCCOM53635B10741,4472,549,009SHSOLE2,117,675 0 431,334
LITHIA MTRS INCCOM53679710313,20561,546SHSOLE59,329 0 2,217
LITTELFUSE INCCOM5370081044,27021,491SHSOLE21,491 0 0
LIVEPERSON INCCOM53814610111,2511,194,403SHSOLE966,423 0 227,980
LOVESAC COMPANYCOM54738L10925,4881,250,655SHSOLE1,040,314 0 210,341
LUXFER HLDGS PLCSHSG5698W1161,985136,907SHSOLE136,907 0 0
MAGNITE INCCOM55955D10068,44610,417,962SHSOLE9,552,110 0 865,852
MATCH GROUP INC NEWCOM57667L1073,06264,130SHSOLE64,130 0 0
MATERION CORPCOM57669010113,903173,784SHSOLE149,306 0 24,478
MAXCYTE INCCOM57777K10663898,201SHSOLE98,201 0 0
MESA LABS INCCOM59064R1094,91434,895SHSOLE29,598 0 5,297
MONOLITHIC PWR SYS INCCOM6098391053,2448,927SHSOLE7,071 0 1,856
MOTORCAR PTS AMER INCCOM62007110018312,050SHSOLE12,050 0 0
NATERA INCCOM63230710414,924340,571SHSOLE310,965 0 29,606
NUTANIX INCCL A67059N1086,596316,663SHSOLE256,715 0 59,948
OLINK HLDG ABSPONSORED ADS6807101009,086748,396SHSOLE634,793 0 113,603
ONESPAWORLD HOLDINGS LIMITEDCOMP7368411312,6921,510,907SHSOLE1,252,138 0 258,769
ONTO INNOVATION INCCOM6833441055,93892,707SHSOLE72,826 0 19,881
ORGANOGENESIS HLDGS INCCOM68621F1028124,973SHSOLE24,973 0 0
ORTHOPEDIATRICS CORPCOM68752L10017,496379,193SHSOLE321,989 0 57,204
OSI SYSTEMS INCCOM6710441058,190113,649SHSOLE92,972 0 20,677
PACIFIC BIOSCIENCES CALIF INCOM69404D1083,333574,189SHSOLE461,642 0 112,547
PAR TECHNOLOGY CORPCOM6988841035,315180,000SHSOLE170,690 0 9,310
PAYCOM SOFTWARE INCCOM70432V102118,389358,765SHSOLE295,997 0 62,768
PAYLOCITY HLDG CORPCOM70438V10647,768197,733SHSOLE164,080 0 33,653
PERFICIENT INCCOM71375U10112,192187,507SHSOLE152,091 0 35,416
PHREESIA INCCOM71944F10612,163477,347SHSOLE386,150 0 91,197
PLIANT THERAPEUTICS INCCOM7291391057,961381,093SHSOLE326,145 0 54,948
PORCH GROUP INCCOM73324510429,23212,991,796SHSOLE11,463,583 0 1,528,213
PORTILLOS INCCOM CL A73642K1068,664439,995SHSOLE386,853 0 53,142
POWER INTEGRATIONS INCCOM73927610311,444177,917SHSOLE144,779 0 33,138
PROS HOLDINGS INCCOM74346Y10317,500708,518SHSOLE595,443 0 113,075
PROTHENA CORP PLCSHSG728001088,555141,108SHSOLE120,437 0 20,671
PTC INCCOM69370C1006,59963,087SHSOLE49,655 0 13,432
PURE STORAGE INCCL A74624M10218,287668,130SHSOLE584,704 0 83,426
QUALYS INCCOM74758T3037,13651,195SHSOLE40,324 0 10,871
QUANTERIX CORPCOM74766Q10114,0281,272,939SHSOLE1,150,999 0 121,940
RADWARE LTDORDM818731079,727446,389SHSOLE380,983 0 65,406
RAPID7 INCCOM7534221048,634201,261SHSOLE159,635 0 41,626
RBC BEARINGS INCCOM75524B10420,29997,681SHSOLE79,067 0 18,614
REPLIGEN CORPCOM75991610912,59767,322SHSOLE55,618 0 11,704
RESEARCH SOLUTIONS INCCOM7610251054321,040SHSOLE21,040 0 0
REVANCE THERAPEUTICS INCCOM7613301096,613244,917SHSOLE200,737 0 44,180
RUSH ENTERPRISES INCCL A7818462091,70538,865SHSOLE38,865 0 0
SHOCKWAVE MED INCCOM82489T1045,46519,653SHSOLE16,013 0 3,640
SHOTSPOTTER INCCOM82536T10717,365603,794SHSOLE501,569 0 102,225
SI-BONE INCCOM82570410915,652896,475SHSOLE774,228 0 122,247
SILICON LABORATORIES INCCOM8269191024,13233,475SHSOLE26,372 0 7,103
SILK RD MED INCCOM82710M1002,46754,828SHSOLE54,828 0 0
SILVERGATE CAP CORPCL A82837P40820,614273,574SHSOLE244,615 0 28,959
SITIME CORPCOM82982T1061,37017,405SHSOLE13,685 0 3,720
SNOW LAKE RES LTDCOM83336J208686389,822SHSOLE359,636 0 30,186
SPROUT SOCIAL INCCOM CL A85209W10920,277334,168SHSOLE274,431 0 59,737
SPS COMM INCCOM78463M107107,970869,116SHSOLE712,218 0 156,898
STOKE THERAPEUTICS INCCOM86150R1073,326259,014SHSOLE227,330 0 31,684
STONERIDGE INCCOM86183P10211,980706,769SHSOLE674,776 0 31,993
STRATASYS LTDSHSM8554810125,3181,756,972SHSOLE1,409,178 0 347,794
TECHTARGET INCCOM87874R1007,598128,348SHSOLE101,128 0 27,220
TEXAS CAP BANCSHARES INCCOM88224Q10723,550398,953SHSOLE331,918 0 67,035
THE BEAUTY HEALTH COMPANYCOM CL A88331L10812,1231,028,285SHSOLE870,611 0 157,674
THREDUP INCCL A88556E1023,0841,676,193SHSOLE1,370,417 0 305,776
TOAST INCCL A88878710879,7894,772,099SHSOLE3,895,373 0 876,726
TOPGOLF CALLAWAY BRANDS CORPCOM13119310414,321743,569SHSOLE604,506 0 139,063
VARONIS SYS INCCOM92228010212,317464,452SHSOLE424,886 0 39,566
VEECO INSTRS INC DELCOM92241710015,931869,588SHSOLE729,309 0 140,279
VERACYTE INCCOM92337F10731,8971,921,506SHSOLE1,786,638 0 134,868
VERICEL CORPCOM92346J10812,571541,870SHSOLE462,579 0 79,291
VERITONE INCCOM92347M1001,331236,416SHSOLE186,359 0 50,057
VICOR CORPCOM92581510212,523211,748SHSOLE191,471 0 20,277
VICTORY CAP HLDGS INCCOM CL A92645B10325,6531,100,514SHSOLE909,975 0 190,539
VITA COCO CO INCCOM92846Q10724,6572,164,836SHSOLE2,061,338 0 103,498
VIZIO HLDG CORPCL A COM92858V1012,962338,912SHSOLE312,254 0 26,658
WESTERN DIGITAL CORP.COM95810210575023,042SHSOLE18,124 0 4,918
WILLSCOT MOBIL MINI HLDNG COCOM CL A97137810443,8931,088,340SHSOLE932,030 0 156,310
WNS HLDGS LTDSPON ADR92932M10110,114123,582SHSOLE99,430 0 24,152
WORKIVA INCCOM CL A98139A10546,204593,882SHSOLE490,702 0 103,180
XENCOR INCCOM98401F1054,373168,307SHSOLE143,438 0 24,869
XENON PHARMACEUTICALS INCCOM98420N10510,825299,855SHSOLE251,075 0 48,780
ZETA GLOBAL HOLDINGS CORPCL A98956A10547,0487,117,707SHSOLE5,904,168 0 1,213,539
ZYMEWORKS INCCOM98985W1023,615587,808SHSOLE505,814 0 81,994